WADDLING WITH PTEROSAURS: NEW PTERAICHNUS TRACKS FROM THE UPPER SUNDANCE FORMATION OF WYOMING
Over 150 pterosaur tracks are preserved as impressions and natural casts in multiple layers of fine-grained, ripple-marked, quartz sandstones in the Late Jurassic (Late Oxfordian-Early Kimmeridgian) Windy Hill Sandstone Member of the Upper Sundance Formation. Preserved in the sediments of ancient beach sands, the plantigrade, v-shaped pes prints typically display four digits, while tracks of the digitigrade manus show two short digits and one long, wing-bearing digit. Pes lengths vary from 6 to 12.8 cm. Pes widths vary from 1.8 to 4.5 cm. The deeper-impressed, manus prints range in length from 4 to 12 cm, with widths ranging from 1.8 to 5.9 cm. Most tracks compare favorably to the ichnospecies, Pteraichnus stokesi, as they are morphologically similar, found within the same stratigraphic level, and fall within the same size range. Over twelve trackways have been located, with two to six sets of pes and manus tracks preserved. The trackways are generally oriented to the northwest and northeast. Trackways display outwardly rotated, manus prints lateral to and behind the pes prints. Most trackways suggest animals of different sizes moving by quadrupedal motion in a linear direction, while track impressions higher in the section demonstrate a pattern of trampling.