Paper No. 260-6
Presentation Time: 2:55 PM
AN EARLY UPLAND LAGERSTÄTTE FROM THE PENNSYLVANIAN (BASHKIRIAN) NARRAGANSETT BASIN OF MASSACHUSETTS, USA
The Carboniferous is well known for vast lowland swamps that provided the taphonomic conditions for preservation of abundant fossils. Little is known, however, of upland ecosystems and niches, which are seldom preserved due to the low preservational potential of xeric to sub-xeric settings and geologic reworking. Upland flora and fauna are often only recorded as allochthonous deposits in coastal plain depositional settings. We report the discovery of a fully terrestrial, upland ichnofossil Lagerstätte from the Lower Pennsylvanian (Bashkirian) of Massachusetts, USA. Wamsutta Fm. red beds consist of thick cross-bedded sandstone and tabular conglomerate units and discontinuous, thin sequences of centimeter-scale laminated fine sandstone and shale, indicative of distal fan and braidplain environments. Prior floral analysis indicated an Early Pennsylvanian age (~318.5 - 317.0 Ma; Bashkirian) but more recent work suggested a mix of early and mid-Pennsylvanian flora. Fossils recovered from the site include diverse vertebrate tracks and trackways with and without tail drags (amphibians, non-mammalian synapsids, and eureptiles), ~130 species of plant morphotaxa that include in situ tree stumps and evidence of insect damage, megaspores, fungal rhizomorphs, ~2 dozen different ichnospecies of full- and partial-body impressions of living insects representing both pterygotes and apterygotes, including the first fossil examples of vertebrate-insect predation, arachnid full-body impressions (thelyphonid) made in life and while molting (ricinuleid), body impressions and trails of myriapods, limulids, and swimming trails (Undichna and Lunichnium), and other invertebrate ichnotaxa (tracks and trails) including new ichnospecies. The site is temporally and ecologically valuable given the abundance of identifiable interactions among species, and the exceptional preservation of the fossils provide an unparalleled glimpse into Carboniferous upland environments.