PHYLOGENETIC REVISION OF THE ORDOVICIAN BRACHIOPOD GENUS GLYPTORTHIS
Type specimens from museum collections were examined to determine character states for phylogenetic analysis of all North American species of Glyptorthis. Non-type and specimens figured in the literature were also examined when possible to assess the range of morphologic variation within each species. A phylogenetic analysis was performed on both internal and external morphological characters using parsimony-based PAUP software and analyzed with MacClade. Both discrete and continuous characters were used. Continuous characters states were differentiated through morphometric analysis and coded as qualitative states.
A set of well supported trees were recovered from parsimony analysis. The strict consensus of these is highly resolved and delineates four monophyletic clades within the genus. Each clade is supported by multiple synapomorphies. These relate primarily to characters of shell ornamentation, valve convexity, shape of the ventral muscle field, and angular relationships within the ventral cardinal area. Preliminary biogeographic analysis suggests that G. insculpta invaded the Cincinnati region from basins in either the central or southeast USA.